B.A., Earlham College
M.A., Harvard University
Ph.D., University of California at Davis

Research Interests

-Behavior and web structure of orb-weaving spiders.

Behavior and physiology research includes work of orb-weaving spiders, dragonflies, baboons, and unusual animal behavior before earthquakes.

-How undergraduates learn to read scientific papers.

-Undergraduate biology teaching and curriculum issues.